Corresponding author: Alexey S. Sazhnev (
Academic editor: R. Yakovlev
The recent status of the family
Sazhnev AS, Dragan SV (2020) New data on the fauna of Heteroceridae (Coleoptera) of the Republic of Khakassia (Eastern Siberia). Acta Biologica Sibirica 6: 399–406
The variegated mud-loving beetles (
According to modern catalogues and articles, the fauna of
In recent publications, the information about
There are ten species of the
The variegated mud-loving beetles were collected in the upper part of the Yenisei River basin (Republic of Khakassia, Abakan city, Abakan River valley, Figs
Second author collected the majority of examined specimens; all specimens are deposited in collection of the Papanin Institute for Biology of Inland Waters of the Russian Academy of Sciences (IBIW, Borok, Yaroslavl Region, Russia; depository and collector not listed in the text). Map was created in SimpleMappr online service (
Collecting site of the variegated mud-loving beetles in the Abakan River valley (Abakan city,
Sampling localities: squares - Shirinsky distr., Belyo Lake; circles - Abakan, Abakan River valley.
The species is known from Asia: China (Shaanxi) (
It is a widespread Holarctic species, also known from the Oriental region: Laos, Philippines, Thailand, Vietnam (
This species distributed in Europe, Turkey, Iran, Kazakhstan, Tajikistan, southern part of Western and Eastern Siberia (
The species is known from Europe, Turkey, Iran, Central Asia (Kazakhstan, Kyrgyzstan, Turkmenistan, Uzbekistan), south of Siberia to Cisbaikalia (
It is known from Europe, Southwest (partly) and Iran, Central Asia (Kazakhstan, Kyrgyzstan, Tajikistan, Uzbekistan), Mongolia, south of Western and Eastern Siberia (
This species distributed in the southern part of Europe, Turkey, Iran, Lebanon, Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, Mongolia, south of Western and Eastern Siberia (
A. Sazhnev was partially supported by the Russian State Assignment (aaaa-a18-118012690105-0). The authors declare no competing interests.